"Sexual Reproduction in Flowering Plants: Processes and Significance"
Flowers are the reproductive structures of angiosperms that contain both male and female reproductive organs. The male reproductive organ is called the androecium and is made up of stamens, while the female reproductive organ is called the gynoecium and consists of pistils.
The stamen has an anther, which contains microsporangia that produce pollen grains. The microsporangia are surrounded by four wall layers, including the epidermis, endothecium, middle layers, and the tapetum. The sporogenous tissue in the centre of the microsporangia undergoes meiosis to produce tetrads of microspores that mature into pollen grains. The pollen grains have a two-layered wall, the outer exine and inner intine. The exine is made up of sporopollenin and has germ pores. Pollen grains may have two cells (a vegetative cell and generative cell) or three cells (a vegetative cell and two male gametes) at the time of shedding.
The pistil has three parts, the stigma, style, and ovary. The ovules are present in the ovary, and each ovule has a stalk called the funicle, protective integuments, and an opening called the micropyle. The nucellus is the central tissue in which the archesporium differentiates. The megaspore mother cell divides meiotically, and one of the megaspores forms the embryo sac, which is the female gametophyte. The mature embryo sac is 7-celled and 8-nucleate, with the egg apparatus at the micropylar end, three antipodals at the chalazal end, and a large central cell with two polar nuclei in the centre.
Pollination is the process by which pollen grains are transferred from the anther to the stigma. Pollinating agents can be either abiotic, such as wind and water, or biotic, such as animals. Pollen-pistil interaction involves events from the landing of pollen grains on the stigma until the pollen tube enters the embryo sac. Following compatible pollination, the pollen grain germinates on the stigma, and the resulting pollen tube grows through the style, enters the ovules, and finally discharges two male gametes in one of the synergids. Angiosperms exhibit double fertilisation because two fusion events occur in each embryo sac, namely syngamy and triple fusion. The products of these fusions are the diploid zygote and the triploid primary endosperm nucleus (in the primary endosperm cell).
The developing embryo passes through different stages, such as the proembryo, globular, and heart-shaped stages, before maturation. The mature dicotyledonous embryo has two cotyledons and an embryonal axis with epicotyl and hypocotyl, while the embryo of monocotyledons has a single cotyledon. After fertilisation, the ovary develops into a fruit, and the ovules develop into seeds.
Some angiosperms exhibit a phenomenon called apomixis, which results in the formation of seeds without fertilisation. Apomicts have several advantages in horticulture and agriculture. Some angiosperms produce more than one embryo in their seed, which is called polyembryony.
In conclusion, the process of sexual reproduction in angiosperms involves the development of male and female reproductive structures, pollination, fertilisation, and the development of fruits and seeds. The process is intricate and involves several stages that ensure the continuation of the species. The phenomenon of apomixis and polyembryony is also essential to understand in the context of angiosperm reproduction.
# Mind Map
- Flower and Pre-Fertilisation Events
- Flower
- Pre-fertilisation:Structures and Events
- Pollination
- Artificial Hybridisation
- Double Fertilisation
- Post Fertilisation : Structure and Events
- Development of an Endosperm
- Development of an Embryo
- Development of a Seed
- Seed Dispersal
- Formation of a Fruit
- Some Special mechanisms of Reproduction
=>Microsporogenesis:
~Microsporogenesis is the process of formation of microspores, which give rise to the male gametophyte (pollen).
~It occurs in the anther of the flower.
~Microsporogenesis involves meiosis, a type of cell division that results in the formation of four haploid microspores from a diploid microsporocyte.
~The four haploid microspores are genetically different from each other and from the parent cell.
~Each microspore undergoes mitosis to form a two-celled pollen grain, which contains the generative cell and the tube cell.
~At the end of microsporogenesis, four haploid microspores are formed.
~Megasporogenesis is the process of formation of megaspores, which give rise to the female gametophyte (embryo sac).
~It occurs in the ovule of the flower.
~Megasporogenesis involves meiosis, a type of cell division that results in the formation of four haploid megaspores from a diploid megasporocyte.
~However, in most cases, only one of the four megaspores survives and develops into the female gametophyte.
~The female gametophyte is usually seven-celled and eight-nucleate.
~At the end of megasporogenesis, one haploid megaspore is formed.
At the end of megasporogenesis, one haploid megaspore is formed, which will eventually develop into a female gametophyte (embryo sac) containing seven cells and eight nuclei.
~Sporogenous tissue - The cells in the sporogenous tissue undergo meiosis to produce microspore mother cells.
~Pollen mother cell - The microspore mother cell undergoes meiosis to produce a tetrad of four haploid microspores.
~Microspore tetrad - The microspore tetrad then separates, and each microspore develops into a pollen grain.
~Pollen grain - The pollen grain is a two-celled structure consisting of a generative cell and a tube cell.
~Male gametes - The generative cell divides to form two haploid male gametes, which are contained within the pollen grain.
The ovule is a reproductive structure found inside the ovary of a flower. It consists of several distinct parts:
~Micropyle: A small opening in the integument that allows for the entry of pollen.
Integument: One or more layers of protective tissue that surround the nucellus.
~Nucellus: The innermost layer of the ovule, which contains the megaspore mother cell.
Megaspore mother cell: The cell within the nucellus that undergoes meiosis to produce a single functional megaspore.
~Embryo sac: The female gametophyte, which is formed by the mitotic division of the functional megaspore. It contains seven cells and eight nuclei.
~Polar nuclei: Two nuclei located in the center of the embryo sac.
~Egg cell: A single haploid cell located at the end of the embryo sac closest to the micropyle. It is the cell that is fertilized by a sperm cell to produce the zygote.
Q 5 . What is meant by monosporic development of female gametophyte?
~The chalazal end: This end of the embryo sac contains three antipodal cells, which are haploid and usually non-functional.
~The micropylar end: This end of the embryo sac contains two synergids, which flank the egg cell, also known as the oosphere. The synergids have filiform apparatus that help guide the pollen tube to the egg cell.
~The central cell: This cell contains two polar nuclei, which are haploid and fuse to form the diploid secondary nucleus. The secondary nucleus is involved in the process of double fertilization.
~The egg cell (1n) is located at the micropylar end of the embryo sac, flanked by two synergids.
~The central cell (2n) is located in the center of the embryo sac, containing two polar nuclei that fuse to form the secondary nucleus (2n).
~Three antipodal cells (1n) are located at the chalazal end of the embryo sac.
~Two polar nuclei (1n) are located within the central cell, and fuse to form the secondary nucleus (2n).
The arrangement of the cells and nuclei is crucial for the process of double fertilization, which involves the fusion of one sperm cell with the egg cell to form the zygote, and the fusion of the other sperm cell with the secondary nucleus to form the triploid endosperm.
Cleistogamous flowers, on the other hand, are self-pollinating flowers that remain closed and do not open. These types of flowers do not rely on pollinators and are adapted for self-pollination, which occurs when the pollen from the anther is transferred directly to the stigma of the same flower.
While cross-pollination can occur in cleistogamous flowers, it is less likely than in chasmogamous flowers. This is because the closed structure of cleistogamous flowers makes it difficult for pollinators to access their reproductive organs and transfer pollen between different flowers. However, some species of plants with cleistogamous flowers have mechanisms to ensure cross-pollination, such as the formation of specialized structures to facilitate the transfer of pollen.
Q 8 . Mention two strategies evolved to prevent self-pollination in flowers.
~Dichogamy: In this strategy, the stamens and pistils of a flower mature at different times, preventing self-pollination. There are two types of dichogamy: protandry and protogyny. In protandry, the stamens mature before the pistils, whereas in protogyny, the pistils mature before the stamens.
~Herkogamy: In this strategy, the physical arrangement of the stamens and pistil in a flower prevents self-pollination. For example, in some flowers, the stamens and pistil are positioned in a way that the pollen from the stamens can only come in contact with the stigma of another flower, and not with its own stigma. Another example is the presence of specialized structures, such as a long style or a narrow corolla tube, which prevent self-pollination by making it difficult or impossible for the pollen to reach the stigma of the same flower.
In self-incompatible species, the female reproductive organ of the flower, the pistil, has the ability to recognize its own pollen, and triggers a rejection mechanism to prevent self-fertilization. This rejection can occur at different stages, including the inhibition of pollen germination on the stigma or the arrest of pollen tube growth within the style.
Self-pollination in self-incompatible species does not lead to seed formation because the plant has evolved mechanisms to prevent fertilization by its own pollen. If self-pollination occurs, the female reproductive system will recognize the pollen as self and trigger the rejection mechanism, leading to the failure of fertilization. As a result, the plant must rely on cross-pollination to reproduce successfully. This promotes genetic diversity and helps to prevent the accumulation of harmful mutations.
Q 10 . What is bagging technique? How is it useful in a plant breeding programme?
The bagging technique is particularly useful in the breeding of crops with specific traits, as it allows breeders to ensure that only desired crosses take place. This helps in maintaining genetic purity of the offspring, which can be important for the success of the breeding program. Additionally, bagging can be used to protect flowers from pests and diseases, and to prevent hybridization with related species that may occur in the area.
The nuclei involved in triple fusion are the two polar nuclei of the central cell and one of the two sperm nuclei from the pollen tube. The other sperm nucleus fuses with the egg cell in a process called syngamy, resulting in the formation of a diploid (2n) zygote. Both triple fusion and syngamy occur after successful pollination and the growth of the pollen tube through the style of the flower.
Q 12 . Why do you think the zygote is dormant for sometime in a fertilised ovule?
Q 14 . Why is apple called a false fruit? Which part(s) of the flower forms the fruit?
In general, the fruit of a flowering plant develops from the ovary of the flower, which encloses and protects the developing seeds. The ovary wall may thicken and become fleshy or hard, forming what we commonly recognize as the fruit. Therefore, the fruit is primarily formed from the ovary of the flower, although other parts of the flower such as the receptacle may contribute to the fruit's structure.
Q 15 . What is meant by emasculation? When and why does a plant breeder employ this technique?
Plant breeders employ this technique when they want to create hybrid seeds that carry desirable traits of two different varieties of the same species. Emasculation ensures that the pollen used in pollination comes from the desired male parent. For example, if a breeder wants to create a hybrid between two tomato varieties, they may emasculate one variety and use the pollen from the other to cross-pollinate the emasculated flower. This ensures that the resulting seeds will carry the desired traits of the second variety.
Q 16 . If one can induce parthenocarpy through the application of growth substances, which fruits would you select to induce parthenocarpy and why?
Fruits such as banana, grape, and pineapple are commonly induced for parthenocarpy because seedless varieties of these fruits are more commercially valuable. For example, seedless grapes and bananas are preferred by consumers because they are easier to eat and can be used in a variety of culinary applications. Additionally, seedless varieties of these fruits are easier to propagate and can be more consistent in their quality and yield. Therefore, inducing parthenocarpy can be a useful technique for plant breeders and growers to produce high-quality fruits that are in demand in the market.
Q 17 . Explain the role of tapetum in the formation of pollen-grain wall.
The tapetum is responsible for synthesizing and depositing the sporopollenin, which is a major component of the pollen wall. Sporopollenin is a highly resistant and durable polymer that provides mechanical and chemical protection to the developing pollen grain. The tapetum also produces the precursors of pollen exine, such as fatty acids and carotenoids, which are transported to the developing microspores and incorporated into the pollen wall.
In addition to its nutritive function, the tapetum also plays a crucial role in the regulation of the timing and pattern of pollen development. It controls the timing of meiosis and the onset of microsporogenesis, and influences the spatial arrangement of the developing microspores. The tapetum also helps to maintain a suitable environment for pollen development by regulating the pH, osmotic potential, and water content of the developing anther.
Overall, the tapetum is essential for the proper development and maturation of pollen grains, and its dysfunction can lead to male sterility in plants.
Apomixis is important in agriculture and plant breeding as it offers the potential to create clones of plants with desirable traits without relying on traditional methods of plant breeding, which can be time-consuming and unpredictable. Apomixis can also help in maintaining heterozygosity of a hybrid variety over generations, allowing for stable production of high-yielding crops. Moreover, apomixis can preserve rare or endangered plant species and facilitate the production of new cultivars that can be used for commercial production.
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FAQs on Sexual Reproduction in Flowering Plants
What is sexual reproduction in flowering plants?
Sexual reproduction in flowering plants is the process by which male and female reproductive structures combine to create offspring with genetic diversity.
What are the reproductive structures of flowering plants?
The reproductive structures of flowering plants include the stamen (male) and the pistil (female).
How is pollen produced in flowering plants?
Pollen is produced in the anthers of the stamen through a process called meiosis.
What is pollination in flowering plants?
Pollination in flowering plants is the process by which pollen is transferred from the anther to the stigma, leading to fertilization and seed formation.
What are the different mechanisms of pollination in flowering plants?
The different mechanisms of pollination in flowering plants include self-pollination, cross-pollination, and animal-mediated pollination.
What is double fertilization in flowering plants?
Double fertilization is a process unique to flowering plants in which two sperm cells from a pollen grain fertilize two different cells within the female reproductive structure of a flower, resulting in the formation of both a zygote and endosperm.
How many sperm cells are involved in double fertilization in flowering plants?
Two sperm cells are involved in double fertilization in flowering plants.
What is the role of the endosperm in flowering plant reproduction?
The endosperm is a nutritive tissue that forms within the seed of a flowering plant and provides nourishment to the developing embryo. It is formed as a result of double fertilization, where one sperm cell fuses with the egg cell to form the zygote, and the other sperm cell fuses with the two polar nuclei to form the endosperm nucleus.
How does the embryo develop in flowering plants?
The embryo in flowering plants develops from the zygote, which is formed when a sperm cell fuses with an egg cell during fertilization. The zygote undergoes a series of divisions to form an embryo that is eventually surrounded by the endosperm and protective seed coat to form a mature seed.
What is seed formation in flowering plants?
Seed formation in flowering plants is the process by which a fertilized ovule develops into a mature seed. It involves the growth and differentiation of the embryo and endosperm, as well as the development of a protective seed coat around the embryo.
How are seeds dispersed in flowering plants?
Seeds in flowering plants can be dispersed by various means such as wind, water, animals, and explosive mechanisms.
What is the significance of sexual reproduction in flowering plants?
Sexual reproduction in flowering plants promotes genetic diversity, which increases the adaptability of the species to changing environments and enhances their chances of survival and evolution.
What are the advantages of sexual reproduction over asexual reproduction in flowering plants?
Sexual reproduction in flowering plants results in genetically diverse offspring, which can better adapt to changing environments and have increased resistance to pests and diseases. Asexual reproduction, on the other hand, produces genetically identical offspring that are more susceptible to these environmental factors.
What is the role of flowers in sexual reproduction in flowering plants?
The primary role of flowers in sexual reproduction is to produce and protect the reproductive structures of the plant, which include the stamen and pistil. The flower attracts pollinators, such as insects and birds, which aid in the transfer of pollen between flowers of the same species, leading to fertilization and seed production.
What is the function of the stamen in sexual reproduction in flowering plants?
The stamen is the male reproductive structure in flowering plants, which consists of the filament and anther. The anther produces pollen, which contains the male gametes, and the filament supports the anther. The stamen plays a critical role in sexual reproduction by producing and delivering pollen to the female reproductive structure, the pistil, for fertilization.
How do the structures of the calyx and corolla protect the reproductive organs in flowering plants?
The calyx and corolla are the outermost parts of the flower. The calyx is typically green and consists of sepals, while the corolla is usually brightly colored and made up of petals. These structures protect the reproductive organs of the flower from external damage and also attract pollinators. Sepals and petals are modified leaves, and their shapes and colors have evolved to suit the specific pollinators that the plant wants to attract.
What is the gynoecium in flowering plants?
The gynoecium is the female reproductive part of the flower. It consists of one or more carpels, which each contain an ovary, style, and stigma. The ovary contains the ovules, which are the female gametes that are fertilized by the male gametes from the pollen. The style is a stalk-like structure that connects the ovary to the stigma, and the stigma is the sticky surface where the pollen lands and germinates.
What is the androecium in flowering plants?
The androecium is the male reproductive structure of a flowering plant, consisting of the stamens, each of which typically contains a filament and an anther. The anther produces and releases pollen, which contains the male gametes.
How does the size and shape of a flower affect pollination in flowering plants?
The size and shape of a flower can affect pollination in different ways. For example, flowers with a long and narrow corolla tube are often pollinated by insects with long tongues, while flowers with a wide and open corolla are often visited by a variety of insects. Flowers with bright colors and strong scents are usually more attractive to pollinators than dull-colored and unscented flowers.
How does sexual reproduction in flowering plants contribute to genetic diversity?
Sexual reproduction in flowering plants contributes to genetic diversity by combining genetic information from two different individuals (the male and female parents) to produce a unique offspring with a new combination of traits. This variation allows for adaptation to changing environments and increases the chances of survival and reproduction of the species.
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